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The Cercopithecinae subfamily comprises all the non-Colobinae African monkeys and is further subdivided into two main tribes: Papionini or baboon-macaques and Cercopithecini or guenons.All major SIV lineages known to date were discovered because their primate hosts had antibodies that cross-reacted with HIV-1 or HIV-2 antigens (10, 11, 15, 19, 22, 26, 33, 30, 41, 45, 72, 74, 95).The presence of a vpu gene has now also been documented in SIVs from several Cercopithecus species: in SIVgsn from a greater spot-nosed monkey (C.nictitans) in Cameroon (27), in SIVmon and SIVmus from mona (C.mona) and mustached (C.cephus) monkeys respectively, and SIVwol from a Wolf's monkey (C.wolfi) from the Democratic Republic of Congo (24, 32).The divergence among Vpu proteins is extremely high, less than 30% identity (27).The only SIVs actually known to have been transmitted to humans, carried either a vpx or a vpu gene.

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